Swifty the Arctic Fox (Jeremy Renner) works in the mail room of the Arctic Blast Delivery Service, but he has much bigger dreams. He yearns to become a Top Dog, the Arctic's star husky couriers. To prove he can do it, he commandeers one of the sleds and delivers a mysterious package to a secret location. Once there, he stumbles on a hidden fortress overseen by the nefarious Otto Von Walrus (John Cleese). The blubbery evil genius commands an army of oddly polite puffin henchmen. Swifty discovers Otto Von Walrus' villainous plan to drill beneath the snow-packed surface to unleash masses of ancient gas to melt the Arctic and become the world's supreme ruler. To stop this sinister scheme, Swifty enlists the help of his friends: PB (Alec Baldwin), a neurotic polar bear, Lemmy (James Franco), a scatterbrained albatross, Jade Fox (Heidi Klum), a brainy engineer, Leopold (Omar Sy) and Bertha (Heidi Klum), two conspiracy theorist otters and Magda (Anjelica Huston), his curmudgeonly moose boss.